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humanity was decimated by the epidemic of Gastrea, a parasitic virus, and is forced to live inside walls, which are created from Varanium: a metal that is capable of subduing Gastrea. Soon, the 'cursed children', children born with the Gastrea virus who are able to control it, giving them superhuman abilities are discovered. Because the 'Gastrea' virus could only be seen in women. The Civilians are trained to specialize in fighting Gastrea, operating alongside an initiator and a 'cursed son', and a promoter, who serves to carry them. Ten years after the epidemic, Rentaro Satomi, a high school student who is also a promoter along with his childhood friend Kisara Tendo and his initiator Enju Aihara, is given a secret mission to prevent the destruction of the Tokyo area.